US Secretary of State John Kerry says there is overwhelming evidence of Russian complicity in the downing of a Malaysia Airlines plane in Ukraine.

Mr Kerry called on Russia to take responsibility for the actions of the rebels, saying their handling of the dead had been "grotesque".

All 298 people on flight MH17 died when it was reportedly hit by a missile.

Russia has been accused of providing the rebels with an anti-aircraft system that was allegedly used in the attack.

Earlier, the remains of up to 196 people were loaded on to refrigerated rail wagons in eastern Ukraine, to be taken to an unknown destination.

Australia's Foreign Minister Julie Bishop has called on pro-Russian separatists not to use the bodies as pawns in their conflict with the Ukrainian authorities.

"There are 298 bodies on that site – their families, their loved ones want them home now," she said.

Rescue workers collect bodies of victims at the site of the crash in Grabove - 20 July 2014 Nearly 200 bodies have been moved from the crash site after three days of being left in the open
Railway employees work near refrigerator wagons in Torez carrying the bodies of victims of the plane crash - 20 July 2014 A freight train carrying the bodies is currently standing at Torez station 15km (nine miles) from the crash site

'Wake-up call'

Mr Kerry said the US had seen major military supplies moving into Ukraine from Russia in the last month, including a convoy of armoured personnel carriers, tanks and rocket launchers.

John Kerry: "The lack of access makes its own statement about culpability and responsibility".

Intercepted calls suggested a Russian SA-11 missile system had been transferred to the rebels, Mr Kerry said, and the US had seen a video of a launcher being moved back into Russia after flight MH17 crashed.

"There's [an] enormous amount of evidence that points to the involvement of Russia in providing these systems, training the people on them," Mr Kerry said on a US TV network.

He also threatened further sanctions on Russia and called on European allies to get tougher with President Putin after the "wake-up call".

Writing in a newspaper on Sunday, UK Prime Minister David Cameron said Europe and the West "must fundamentally change our approach to Russia" if Mr Putin "does not change his approach to Ukraine".

A rescue worker inspects the main crash site of the Boeing 777 in Grabove - 20 July 2014 A team of Malaysian experts remain in Kiev after being unable to travel to the crash site in Grabove

At the scene: Fergal Keane, BBC News, at the crash site

The indiscipline and chaos of the last two days have been replaced by the robust presence of former riot policemen who now form a cordon around the central area of the crash site.

There is still no sign of the fully fledged independent investigation which is being demanded by the international community.

During the morning local volunteers have been searching the fields. We have been told that their job is to pinpoint belongings and remains to the emergency services. But this could easily have the effect of disturbing evidence important to an inquiry.

As for the strong words from British Prime Minister David Cameron attacking Russia they are likely to have little impact on the rebels here. They are contemptuous of the West and are much more concerned with the local military balance than with any warnings from London.


Bodies moved

Mr Kerry's comments joined a chorus of anger directed at Russian President Vladimir Putin after the rebels in charge of the crash site restricted access to the plane's wreckage and its victims.

The remains of up to 196 people were loaded on to refrigerated rail wagons at the crash site in Grabove on Sunday, three days after the Boeing 777 came down.

The freight train is now standing at Torez railway station, 15km (nine miles) from the crash site.

Ukrainian workers move debris from the site of the plane crash in eastern Ukraine - 20 July 2014 Heavy machinery has been used to move many of the larger parts of debris from the Boeing 777

The carriages, with heavy closed doors, look like refrigeration units and there is the occasional smell normally associated with dead bodies, the BBC's Richard Galpin reports from Torez.

Speaking in Donetsk, the biggest rebel-held city in the east, rebel leader Alexander Borodai said the bodies would remain in Torez until international aviation inspectors arrived.

He said the bodies had been moved "out of respect for the families", adding: "We couldn't wait any longer because of the heat and also because there are many dogs and wild animals in the area."

'Site compromised'

The rebels say they will hand over MH17's flight recorders to the International Civil Aviation Organization but the US state department said rebels had tampered with other potential evidence.

Heavy machinery could be seen moving plane debris around at the crash site on Sunday.

Footage appears to show one of the plane's data recorders being moved

Malaysian Transport Minister Liow Tiong-Lai, who arrived in Ukraine's capital Kiev at the weekend, said he was "very concerned" that the crash site had been "severely compromised".

A Malaysian team of 133 officials and experts, comprising of search and recovery personnel, forensics experts, technical and medical experts have also arrived in Ukraine.

But the government in Kiev says it has been unable to establish a safe corridor to the crash site for the group.

Fighting remains ongoing in eastern Ukraine between the separatist rebels and government forces in a conflict which erupted in April and is believed to have claimed more than 1,000 lives.

Mourners attend a memorial service in Melbourne held for a family of five killed in the flight MH17 crash - 20 July 2014 Several memorial services were held in Australia on Sunday for victims of the crash in Ukraine

The passenger list released by Malaysia Airlines shows the plane was carrying 193 Dutch nationals, including one with dual US nationality.

Other victims included 43 Malaysians (including 15 crew), 27 Australians, 12 Indonesians, 10 Britons, four Germans, four Belgians, three from the Philippines, and one from both Canada and New Zealand.

Memorial services and vigils were held on Sunday in many countries, including Australia, Malaysia and the Netherlands.